Danny Care and Alex Lozowski have been dropped by England following the laboured 35-15 victory over Japan while Nathan Hughes has been included in England’s training squad for the climax to the autumn series against Australia at Twickenham.
Hughes is available to face the Wallabies after serving his six-week ban for punching and conduct prejudicial to the interest of the game.
The Wasps No 8 has replaced Zach Mercer among the 28 summoned to England’s Surrey training base for the final Test. Piers Francis has been called up to replace Lozowksi.
Richard Wigglesworth, a second-half replacement on Saturday, is poised to supply scrum-half cover for Ben Youngs against Australia after ousting Care.
Lozowski has paid the price for failing to stop centre Ryoto Nakamura scoring Japan’s opening try, a mistake which saw him replaced by Owen Farrell at half-time.
Francis won the most recent of his four caps during the summer tour to South Africa, making a replacement appearance in the first Test at Ellis Park.
Quick guide
England squad
Charlie Ewels (Bath Rugby), Jamie George (Saracens), Dylan Hartley (Northampton Saints), Alec Hepburn (Exeter Chiefs), Ted Hill (Worcester Warriors) *, Nathan Hughes (Wasps), Maro Itoje (Saracens), Courtney Lawes (Northampton Saints), Ben Moon (Exeter Chiefs), Brad Shields (Wasps), Kyle Sinckler (Harlequins), Elliott Stooke (Bath Rugby), Sam Underhill (Bath Rugby), Harry Williams (Exeter Chiefs), Mark Wilson (Newcastle Falcons)
*Apprentice
Chris Ashton (Sale Sharks), Joe Cokanasiga (Bath Rugby), Elliot Daly (Wasps), Owen Farrell (Saracens), George Ford (Leicester Tigers), Piers Francis (Northampton Saints), Jonny May (Leicester Tigers), Jack Nowell (Exeter Chiefs), Henry Slade (Exeter Chiefs), Ben Te’o (Worcester Warriors), Manu Tuilagi (Leicester Tigers), Richard Wigglesworth (Saracens), Ben Youngs (Leicester Tigers)
• More to follow